Study of World Biome
The planet Earth
Biomes are climatically and geographically defined as similar climatic conditions on
the Earth, such as communities of plants, animals, and soil
organisms, and are often referred to as ecosystems. Some parts of the earth have more or less
the same kind of abiotic and biotic factors spread over a large area creating
a typical ecosystem over that area. Such major ecosystems are termed as biomes.
Biomes are defined by factors such as plant structures (such as trees, shrubs, and grasses), leaf types (such as broadleaf and needleleaf), plant spacing (forest, woodland, savanna), and climate. Unlike ecozones, biomes are not defined by genetic,
taxonomic, or historical similarities. Biomes are often identified with
particular patterns of ecological succession and climax
vegetation (quasi-equilibrium
state of the local ecosystem). An ecosystem has many biotopes and a biome is a major habitat type. A major habitat type, however, is a
compromise, as it has an intrinsic inhomogeneity.The biodiversity characteristic of each extinction, especially the diversity of fauna and subdominant plant forms, is a function of abiotic factors and the biomass productivity of the dominant vegetation. In terrestrial biomes, species diversity tends to correlate positively with net primary productivity, moisture availability, and temperature.
Ecoregions are grouped into both biomes and ecozones.
A fundamental classification of biomes is:
- Terrestrial (land) biomes
- Aquatic biomes (including Freshwater
biomes and Marine biomes)
Sometimes an entire biome may be targeted for protection, especially under an individual nation's Biodiversity Action Plan.
Climate is a major factor determining the distribution of terrestrial biomes. Among the important climatic factors are:
- latitude: Arctic, boreal, temperate,
subtropical, tropical.
- humidity: humid, semi-humid, semi-arid, and
arid.
- seasonal variation: Rainfall may be
distributed evenly throughout the year or be marked by seasonal
variations.
- dry summer, wet winter: Most regions
of the earth receive most of their rainfall during the summer months;
Mediterranean climate regions receive their rainfall during the winter
months.
- elevation: Increasing elevation causes a
distribution of habitat types similar to that of increasing latitude.
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Biome Classification Schemes
Biome classification schemes seek to
define biomes using climatic measurements. Particularly in the 1970s and 1980s
there was a significant push to understand the relationships between these
measurements and properties of ecosystem energetics because such discoveries
would enable the prediction of rates of energy capture and transfer among
components within ecosystems. Such a study was conducted by Sims et al. (1978)
on North American grasslands. The study found a positive logistic correlation
between evapotranspiration in mm/yr and above ground net primary
production in g/m^2/yr. More general results from the study were that
precipitation and water use lead to aboveground primary production, solar
radiation and temperature lead to belowground primary production (roots), and
temperature and water lead to cool and warm season growth habit.[3] These findings help explain the categories
used in Holdridge’sbioclassification scheme, which were then later simplified
in Whittaker’s. The number of classification schemes and the variety of
determinants used in those schemes, however, should be taken as a strong indicator
that biomes do not all fit perfectly into the classification schemes created.
Holdridge Scheme
The Holdridge classification scheme
was developed by L. R. Holdridge, a botanist. It maps climates based on four
categories:- Average total precipitation (cm) on a
logarithmic scale
- Potential evapotranspiration ratio:
the potential evapotranspiration divided by the precipitation; the ratio
increases from humid to arid regions.
- Potential evapotranspiration
- Mean annual biotemperature (°C):
calculated from monthly mean temperatures after converting any mean
temperature to 0°C, based on the assumption that temperatures at or below
freezing all have the same effect on plants, and delineating between -10°C
and -30°C would yield unrealistic results.
Whittaker's Biome-type Classification Scheme
Whittaker appreciated biome-types as a representation of the great diversity of
the living world, and saw the need to establish a simple way to classify these
biome-types. Whittaker based his classification scheme on two abiotic factors:
Precipitation and Temperature. His scheme can be seen as a simplification of
Holdridge's, one more readily accessible, but perhaps missing the greater
specificity that Holdrige's provides.Whittaker based his representation of global biomes on both previous theoretical assertions as well as an ever increasing empirical sampling of global ecosystems. Whittaker was in a unique position to make such a holistic assertion as he had previously compiled a review of biome classification.
The Whittaker Classification Scheme can be viewed at the following address:
Key definitions for understanding Whittaker's
Scheme
- physiognomy: The apparent characteristics,
outward features, or appearance of ecological communities or species.
- biome: a grouping of terrestrial ecosystems
on a given continent that are similar in vegetation structure,
physiognomy, features of the environment and characteristics of their
animal communities
- formation: a major kind of community of plants
on a given continent
- biome-type: grouping of convergent biomes or
formations of different continents; defined by physiognomy
- formation-type: grouping of convergent formations
Whittaker's parameters for classifying biome-types
Whittaker, seeing the need for a
simpler way to express the relationship of community structure to the
environment, used what he called “gradient analysis” of ecocline patterns to
relate communities to climate on a worldwide scale. Whittaker considered four
main ecoclines in the terrestrial realm.Intertidal levels: The wetness gradient of areas that are exposed to alternating water and dryness with intensities that vary by location from high to low tide
- climatic moisture gradient
- temperature gradient by altitude
- temperature gradient by latitude
- The gradient runs from favorable to
extreme with corresponding changes in productivity.
- Changes in physiognomic complexity
vary with the favorability of the environment (decreasing community
structure and reduction of stratal differentiation as the environment
becomes less favorable).
- Trends in diversity of structure
follow trends in species diversity; alpha and beta species diversities
decrease from favorable to extreme environments.
- Each growth-form (i.e. grasses, shrubs,
etc.) has its characteristic place of maximum importance along the
ecoclines.
- The same growth forms may be dominant
in similar environments in widely different parts of the world.
Walter System
The Heinrich Walter classification
scheme was developed by Heinrich Walter, a German ecologist. It differs from
both the Whittaker and Holdridge schemes because it takes into account the
seasonality of temperature and precipitation. The system, also based on
precipitation and temperature, finds 9 major biomes, with the important climate
traits and vegetation types summarized in the accompanying table. The
boundaries of each biome correlate to the conditions of moisture and cold
stress that are strong determinants of plant form, and therefore the vegetation
that defines the region. Extreme conditions, such as flooding in a swamp, can
create different kinds of communities within the same biome.- I: Equatorial
- Always moist and lacking temperature
seasonality
- Evergreen tropical rain forest
- II: Tropical
- Summer rainy season and cooler
“winter” dry season
- Seasonal forest, scrub, or savanna
- III: Subtropical
- Highly seasonal, arid climate
- Desert vegetation with considerable
exposed surface
- IV: Mediterranean
- Winter rainy season and summer
drought
- Sclerophyllous (drought-adapted),
frost-sensitive shrublands and woodlands
- V: Warm temperate
- Occasional frost, often with summer
rainfall maximum
- Temperate evergreen forest, somewhat
frost-sensitive
- VI: Nemoral
- Moderate climate with winter freezing
- Frost-resistant, deciduous, temperate
forest
- VII: Continental
- Arid, with warm or hot summers and
cold winters
- Grasslands and temperate deserts
- VIII: Boreal
- Cold temperate with cool summers and
long winters
- Evergreen, frost-hardy needle-leaved
forest (taiga)
- IX: Polar
- Very short, cool summers and long,
very cold winters
- Low, evergreen vegetation, without
trees, growing over permanently frozen soils
Bailey System
Robert G. Bailey almost developed a biogeographical classification system for the United
States in a map published in 1976. Bailey subsequently expanded the system to
include the rest of South America in 1981 and the world in 1989. The Bailey
system is based on climate and is divided into seven domains (Polar, Humid
Temperate, Dry, Human, and Humid Tropical), with further divisions based on
other climate characteristics (subarctic, warm temperate, hot temperate, and
subtropical; marine and continental; lowland and mountain).[7]- 100
Polar Domain
- 120 Tundra Division
- M120 Tundra Division - Mountain
Provinces
- 130 Subarctic Division
- M130 Subarctic Division - Mountain
Provinces
- 200
Humid Temperate Domain
- 210 Warm Continental Division
- M210 Warm Continental Division -
Mountain Provinces
- 220 Hot Continental Division
- M220 Hot Continental Division -
Mountain Provinces
- 230 Subtropical Division
- M230 Subtropical Division - Mountain
Provinces
- 240 Marine Division
- M240 Marine Division - Mountain
Provinces
- 250 Prairie Division
- 260 Mediterranean Division
- M260 Mediterranean Division -
Mountain Provinces
- 300
Dry Domain
- 310 Tropical/Subtropical Steppe
Division
- M310 Tropical/Subtropical Steppe
Division - Mountain Provinces
WWF system
A team of biologists convened by the World Wide Fund for Nature (WWF) developed an ecological land
classification system that
identified fourteen biomes,[8] called major habitat types, and further divided the world's land area
into 867 terrestrial ecoregions. Each terrestrial Ecoregion has a specific
EcoID, fomatXXnnNN (XX is the Ecozone, nn is the Biome number, NN is the individual number). This
classification is used to define the Global
200 list of ecoregions identified by the WWF as priorities for
conservation. The WWF major habitat types are:- 01 Tropical and
subtropical moist broadleaf forests (tropical and subtropical, humid)
- 02 Tropical and
subtropical dry broadleaf forests
(tropical and subtropical, semi-humid)
- 03 Tropical and
subtropical coniferous forests
(tropical and subtropical, semi-humid)
- 04 Temperate broadleaf and
mixed forests (temperate,
humid)
- 05 Temperate coniferous forests (temperate, humid to semi-humid)
- 06 Boreal forests/taiga (subarctic, humid)
- 07 Tropical
and subtropical grasslands, savannas, and shrublands (tropical and subtropical, semi-arid)
- 08 Temperate
grasslands, savannas, and shrublands (temperate, semi-arid)
- 09 Flooded grasslands and savannas (temperate to tropical, fresh or
brackish water inundated)
- 10 Montane grasslands and
shrublands (alpine or montane climate)
- 11 Tundra (Arctic)
- 12 Mediterranean forests,
woodlands, and scrub or Sclerophyll forests (temperate warm, semi-humid to semi-arid with winter rainfall)
- 13 Deserts and xeric shrublands (temperate to tropical, arid)
- 14 Mangrove (subtropical and tropical, salt water
inundated)
Freshwater biomes
According to the World Wildlife Fund,
the following are classified as freshwater biomes:[9]
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Realms or Ecozones (terrestrial and freshwater,
WWF)
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Marine biomes
Marine biomes (H) (major habitat types), Global 200
(WWF)
Biomes of the coastal &continental
shelf areas (Neritic
zone - List of ecoregions (WWF))- Polar
- Temperate shelves and sea
- Temperate upwelling
- Tropical upwelling
- Tropical
coral[10]
Realms or Ecozones (marine, WWF)
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Other marine habitat types
- Hydrothermal vents
- Cold seeps
- Benthic
zone
- Pelagic
zone (trades and westerlies)
- Abyssal
- Hadal (ocean trench)
Major Habitats, Non Global 200 (WWF)
Summary - Ecological taxonomy (WWF)
- Biosphere (List of ecoregions)
- Ecozones or Realms (8)
- Terrestrial Biomes (Major Habitat
Types, 14)
- Ecoregions (867)
- Freshwater Biomes (Major Habitat
Types, 12)
- Ecoregions (426)
- Marine Ecozones or Realms (13)
- Continental Shelf Biomes (Major Habitat Types, 5)
- (Marine
Provinces) (62)
- Ecoregions (232)
- Open & Deep Sea Biomes (Major
Habitat Types)
- Endolithic Biome
- Biosphere
- Ecozone: Palearctic ecozone
- Terrestrial Biome: Temperate Broadleaf and
Mixed Forests
- Ecoregion: Dinaric Mountains mixed forests (PA0418)
- Ecosystem: Orjen, vegetation belt between 1,100-
1,450 m, Oromediterranean zone, Nemoral zone (temperate zone)
- Biotope: Oreoherzogio-AbietetumillyricaeFuk.
(Plant list)
- Plant: Silver fir (Abies
alba)
Anthropogenic biomes
Humans have fundamentally altered
global patterns of biodiversity and ecosystem processes. As a result,
vegetation forms predicted by conventional biome systems are rarely observed
across most of Earth's land surface. Anthropogenic biomes provide an alternative view of the
terrestrial biosphere based on global patterns of sustained direct human
interaction with ecosystems, including agriculture, human settlements,
urbanization, forestry and other uses of land. Anthropogenic biomes offer a new
way forward in ecology and conservation by recognizing the irreversible
coupling of human and ecological systems at global scales and moving us toward
an understanding how best to live in and manage our biosphere and the
anthropogenic biosphere we live in. The main biomes in the world are
freshwater, marine, coniferous, deciduous, ice, mountains, boreal, grasslands,
tundra, and rainforests.
Major Anthropogenic Biomes
- Dense Settlements
- Villages
- Croplands
- Rangelands
- Forested
Other biomes
The Endolithic biome, consisting entirely of microscopic
life in rock pores and cracks, kilometers beneath the surface, has only recently been
discovered and does not fit well into most classification schemes.
Map of Biomes
Freshwater Biomes
Drainage basins of the principal
oceans and seas of the world. Grey areas are endorheic basins that do not drain to the ocean.
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Terrestrial biome
Introduction

Figure 1: Global
distribution of the Earth's thirteen major terrestrial biomes. (Original Data
Source for Map: Olson, D.M. et al. 2001. Terrestrial Ecoregions of the World: A
New Map of Life on Earth. BioScience 51(11): 933-938).
Many places on Earth share similar
climatic conditions despite being found in geographically different areas. As a
result of natural selection, comparable ecosystems have developed in
these separated areas. Scientists call these major ecosystem types biomes. The geographical distribution (and
productivity) of the various biomes is controlled primarily by the climatic
variables precipitation and temperature. The maps in Figures 1 and 2 describe the
geographical locations of the thirteen major terrestrial biomes of the world.
Because of their scale, these maps ignore the many community variations that
are present within each biome category. 
Figure 2:
Distribution of the Earth's thirteen major terrestrial biomes over North and
Central America. (Original Data Source for Map: Olson, D.M. et al. 2001.
Terrestrial Ecoregions of the World: A New Map of Life on Earth. BioScience
51(11): 933-938).
Most of the classified biomes are
identified by the dominant plants found in their communities. For example, the
various types of grasslands are dominated by a variety of annual and perennial
species of grass, while deserts are occupied by plant species that require very
little water for survival or by plants that have specific adaptations to
conserve or acquire water. The diversity of animal life and subdominant plant forms characteristic of each biome is generally controlled by abiotic environmental conditions and the productivity of the dominant vegetation. In general, species diversity becomes higher with increases in net primary productivity, moisture availability, and temperature.
Adaptation and niche specialization are nicely demonstrated in the biome concept. Organisms that fill similar niches in geographically separated but similar ecosystems usually are different species that have undergone similar adaptation independently, in response to similar environmental pressures. The vegetation of California, Chile, South Africa, South Australia, Southern Italy and Greece display similar morphological and physiological characteristics because of convergent evolution. In these areas, the vegetation consists of drought-resistant, hard-leaved, low growing woody shrubs and trees like eucalyptus, olive, juniper, and mimosa.
Tundra
The geographical distribution of the tundra
biome is roughly
poleward of 65° North latitude. In the Southern Hemisphere, the tundra
biome has a very limited
distribution. Within the tundra biome, temperature, precipitation, and
evaporation all tend to be at a minimum. In fact, the tundra is the coldest of
all biomes and this environmental factor has played an important role in the
evolution of adaptations for plant and animal survival. Most tundra locations,
have summer months with an average temperature between 3 and 12° C (37 to 54°
F). The average winter monthly temperature is around -34° C (-30° F).
Precipitation in the wettest month is usually no greater than 2.5 centimeters
(roughly 1 inch). Yet, despite the low levels of precipitation the ground
surface of the tundra biome is often waterlogged because of low rates of
evapotranspiration and poor drainage. The tundra biome is characterized by the absence of trees and the presence of low-lying shrubs, mosses, and lichens. Lack of height allows the vegetation to be protected by the insolating properties of snow during the winter season. Perhaps the most characteristic arctic tundra plants are lichens like reindeer moss (Cladonia spp.). In the drier parts of the tundra, grasses are common (Figure 3). Sedges dominate sites that have more moisture. About 400 varieties of flowering plants occur in this biome. Total species diversity of plants in the tundra biome is relatively small numbering about 2000 species. Plants are generally small, are adapted to soil disturbance, and reproduce via budding or other forms of asexual reproduction rather than sexual means. Soils of this biome are usually permanently frozen (permafrost) starting at a depth of a few centimeters to meter or more. The permafrost line is a physical barrier to plant root growth. Thus, there are no deep rooting systems. The presence of permafrost also causes poor drainage and soils are often waterlogged and chemically reduced.
Figure 3: Tundra dominated by flowering arctic cotton grass, Northwest Territories, Canada. (Image Source).
The principal herbivores of the tundra biome include caribou, musk ox, arctic hare, voles, squirrels, and lemmings (Figure 4). Most of the bird species of the tundra have the ability to migrate and live in warmer locations during the cold winter months. The herbivore species support a small number of carnivore species like the arctic fox, snow owl, polar bear, and wolves. Reptiles and amphibians are few or completely absent because of the extremely cold temperatures.
Alpine tundra is quite comparable to arctic tundra but differs in the absence of permafrost, the presence of better drainage, and more extreme annual fluctuations of air temperature. Plants species in the alpine tundra are for the most part similar to the ones found on the arctic tundra. In contrast, alpine tundra animal species tend to be quit different from those individuals that live in the arctic tundra. This takes place because alpine tundra tends to adopt migrating species during the summer months from habitats located at lower elevations.
Boreal
Forests/Taiga
This moist-cool, transcontinental
boreal forests or taiga biome lies largely between 50 and 65° North
latitude. The climate of this biome is cool to cold with more precipitation
than the tundra. Precipitation here mainly occurs in the summer because this is
the season when mid-latitude cyclones move in from the south. The growth season
is limited to about 130 days. The predominant vegetation of boreal forest biome is cone bearing needle-leaf evergreen variety tree species. Four tree genera are dominant in this biome: spruce (Picea), pine (Pinus), fir (Abies), and larch (Larix). In North America, some common species include: black spruce (Piceamariana), white spruce (Piceaglauca), jack pine (Pinusbanksiana), tamarack (Larixlaricina), and balsam fir (Abiesbalsamea); with red pine (Pinusresinosa), white pine (Pinusstrobus), and hemlock (Tsugacanadensis) limited to an area north and east of the Great Lakes Region. Broad-leaf species, like alder (Alnus), birch (Betula), and aspen (Populus), are common in all areas as an early successional species after disturbance.
Understory vegetation is relatively limited as a result of the low light penetration even during the spring and fall months. Common understory species include orchids, shrubs like rose, blueberry, and cranberry. Mammals common to the boreal forest include moose, bear, deer, wolverine, marten, lynx, wolf, snowshoe hare, vole, chipmunks, shrews, and bats. Reptiles are extremely rare, once again, because of cold temperatures.
Deep litter layers are a common characteristic of boreal forest soils. These deep litter layers accumulate because of slow decomposition rates. Soils of this biome are also acidic and mineral deficient. Mineral deficiency occurs because large amounts of water move down though the profile causing leaching.
Boreal forest soils are characterized by a deep litter layer and slow decomposition. Soils of this biome are also acidic and mineral deficient because of the large movement of water vertically though the profile and subsequent leaching.
Temperate
Coniferous Forests
In North America we can find two
broad areas of temperate coniferous forests in the more temperate
mid-latitudes. In these areas, average annual temperatures range from 20° to 5°
C (68° to 41° F). Along the west side of North America and below the boreal
forest is one such area. On the wetter sites (up to 400 centimeters or 160
inches annually) that have close proximity to the Pacific Ocean are stands of
very tall and productive Douglas fir (Pseudotsugamenziesii), red cedar (Thujaplicata),
sitka spruce (Piceasitchensis), and redwood (Sequoia sempervirens).
Some of these trees can grow to over 120 meters (390 feet) in height. Beneath
the canopy of these trees is a shrub layer that includes various types of
berries (Vacciniumspp.), a few herbs, and various ferns. Further
inland of this temperate “rain forest” zone precipitation declines
significantly, winter temperatures become colder, and summer temperatures
become much warmer. This change in climate makes more drought resistant trees
like ponderosa pine (Pinuspondersoa), Engelmann spruce (Piceaengelmannii), and
lodgepole pine (Pinuscontorta) dominant. Another region of temperature coniferous forests occurs in southeastern United States. The species composition of this forest ecosystem does not resemble the coniferous forests found in western North America. Instead, these forests are dominated by pitch pine (Pinusrigida), longleaf pine (Pinuspalustris), and slash pine (Pinuselliotti). All of these tree species are adapted to growing on nutrient poor sandy soils and can withstand the effects of fire. Biomass productivity is typically low in this type of temperate coniferous forest.
Outside of North America, the various types of temperate coniferous forest can also be found in northern Japan, and parts of Europe and Asia. In these areas, the plant species are similar in form and ecological function to North American species but not closely related.
Temperate Broadleaf and Mixed Forests
The temperature broadleaf and mixed
forests biome (also called temperate deciduous forest) is characterized by a
moderate temperate climate and a dominance of broadleaf deciduous trees. This
biome once occupied much of the eastern half of the United States, central
Europe, Korea, and China. Over the last few centuries, this biome has been very
extensively affected by human activity. Much of it has been converted into
agricultural fields or urban land-use. Tree species diversity is this biome is moderate with 5 to 25 dominant trees at a site. Dominant trees include maple (Acer spp.), beech (Fagus spp.), oak (Quercusspp.), hickory (Caryaspp.), basswood (Tilia spp.), magnolia (Magnolia spp.), cottonwood (Populus spp.), elm (Ulmus spp.), and willow (Salix spp.). The understory of shrubs, herbs, and ferns in a mature forest are typically well developed and richly diversified. Understory plants in this biome often take advantage of the leafless condition of trees during spring and fall to concentrate their growth.
Many different types of herbivores and carnivores live in the temperate broadleaf and mixed forest. Common fauna include squirrels, rabbits, skunks, birds, deer, mountain lion, bobcat, timber wolf, fox, and bears. Some reptiles and amphibians also exist here.
Nutrient rich brown forest soils characterize the temperate broadleaf and mixed forests biome. Tree cover promotes the accumulation of organic materials in a well-developed humus layer. Surface litter layer in these soils tends to be thin because of rapid decomposition.
Temperate Grasslands, Savannas and Shrublands
In central North America is the
temperate grasslands, savannas and shrublands biome (also called prairie). The grassland
biome is also found in the
continental interior of Eurasia, Australia, and South America. Prior to the
arrival of settlers in North America, much of this biome was dominated by
species of tall grass known as bluestem (Andropogon spp.). This particular
species covered much of the eastern side of this biome forming dense covers 1.5
to 2.0 meters (4 to 6 feet) tall. In the western end of the biome, where
precipitation is lower, buffalo grass (Buchloedactyloides) and other
grasses only a few inches above the soil surface are common. Flowering herbs,
including many kinds of composites and legumes, are common but much less
important than grass species. Trees are found scattered in moist low-lying
areas and along a narrow zone adjacent to streams. Climatically, the temperate grasslands, savannas and shrublands biome can be described as being temperate. Summers are hot to warm and winters are cool to cold. Annual precipitation is less than what is received by the adjacent temperate broadleaf and mixed forests biome. Seasonally, precipitation varies from being concentrated during a few months to spread evenly through the year. This biome generally does not receive enough precipitation to support tree growth. In the wetter parts of this biome nutrient rich black chernozemic soils are common. In many parts of the world, these extremely fertile soils now support crop growth. In drier parts of prairies, soils can be influenced by salinization.
Grassland mammals are dominated by smaller burrowing herbivores (prairie dogs, jack rabbits, ground squirrels, and gophers) and larger running herbivores such as bison, pronghorn antelope, and elk. Carnivores include badger, coyote, ferret, wolf, and cougar. The populations of many of these organisms have been drastically reduced because of the conversion of their natural habitat into cropland. Some of these species are on the edge of extinction.
Montane Grasslands and Shrublands
The montane grasslands and shrublands
biome is found at high elevations in temperate, subtropical, and tropical
climates. This biome is dominated by grass and shrub species and tends to have
a high number of endemic plants and animals. Examples of this biome can be
found at the Tibetan plateau, Central Range in New Guinea, eastern Andes
Mountains in South America, southeastern Africa, and tropical East Africa. A
unique feature of many tropical examples of this biome is the presence of giant
rosette vegetation belonging to the plant families Lobelia (Africa), Puya
(South America), Cyathea (New Guinea), and Argyroxiphium (Hawaii) (Figure
7k-16). All of these plants have unique adaptations that allow them to
successfully grow at high elevations.
Deserts
and Xeric Shrublands
In its most typical form, the xeric
shrublands and desert biome consists of shrub-covered land where the
plants are spatially quite dispersed. This biome is geographically found from
25 - 35° North and South latitude, primarily in the interiors of continents.
The formation of precipitation in desert and xeric shrublands biome is limited
by the subtropical high-pressure system. Many desert areas have less than 3
centimeters (about 1 inch) of precipitation during an average year. Dominant plants include drought resistant shrubs like the creosote bush (Larreadivaricata) and sagebrush (Artemisia tridentata), water storing succulents like cactus, and many species of short lived annuals that complete their life cycles during infrequent and short rainy periods (Figure 7k-18). Lastly, desert habitats can be completely devoid of vegetation if precipitation is in very short supply. Most desert mammals tend to be nocturnal to avoid the high temperatures. Desert habitats have a rich lizard and snake fauna because high temperatures promote the success of cold-blooded life forms. Because biomass productivity is low, the litter layer is almost nonexistent and organic content of surface soil layers is very low. Finally, evaporation tends to concentrate salts at the soil surface.
Mediterranean Forests, Woodlands and Scrub
The Mediterranean forests, woodlands
and scrub biome (also called chaparral) has a very specific spatial
distribution. It is found in a narrow zone between 32 and 40° latitude North
and South on the west coasts of the continents. This area has a dry climate
because of the dominance of the subtropical high pressure zone during the fall,
summer, and spring months. Precipitation falls mainly in the winter months
because of the seasonal movement of the polar front and associated mid-latitude
cyclones. Precipitation varies from about 30 to 75 centimeters (12 to 30
inches) annually and most of this rain falls in a period only 2 to 4 months
long. Despite the fact that this biome is very limited geographically, it contains a high diversity of animal and plant species that are adapted to the stressful conditions of long, hot summers with little rain. The vegetation of this biome consists of many different types of annuals and drought-resistant, evergreen, short woody shrubs and trees. Dominant tree species include olive (Oleaeuropaea), eucalyptus (Eucalyptus spp.), arbutus (Arbutus unedo), acacia (Acacia spp.), maritime pine (Pinuspinaster), and various species of oak (Quercus spp.). As a result of the climate, the vegetation of this biome exhibits a number of adaptations to withstand drought and fire. Plants tend not to drop their leaves during the dry season because of the expense of replacement. The dry climate slows the rate of leaf decomposition and soils tend to be poorly developed.
Tropical and Subtropical Grasslands, Savannas and
Shrublands
Vegetation in the tropical and
subtropical grasslands, savannas and shrublands biome (also called savanna)
consists of a cover of perennial grass species 1 to 2 meters (3 to 6 feet) tall
with scattered drought-resistant trees that generally do not exceed 10 meters
(32 feet) in height. The savanna biome constitutes extensive areas in eastern
Africa, South America, and Australia. Distinct wet and dry seasons and
temperatures that are hot all year long characterize the climate of this biome.
Annual rainfall varies between 90-150 centimeters (35 to 60 inches). Tree and shrub species in the savanna usually drop their leaves during the dry season. This adaptation reduces water loss from the plants during the dry winter season. Diversity of plant and animal species tends to be high. Grazing on the grasses and trees are vast herds of hoofed mammals including buffalo, giraffes, eland, impalas, oryx, gazelles, gerenuk, wildebeest, zebra, rhinoceroses, elephants, and warthogs. These herbivores supply food for carnivores like lions, cheetahs, leopards, jackals, and hyenas.
Flooded Grasslands and Savannas
In the tropical and subtropical
regions of our planet are large expanses of flooded grasslands and savannas.
This biome is slightly different from the savanna biome just described. Because
of common flooding, these areas support additional plant and animal species
adapted to thrive under this condition. For instance, this biome is home to
large numbers of migratory and resident water birds. Some examples of flooded grasslands and savannas include in the Everglades in Florida, the Sahelian flooded savannas, and the Zambezian flooded savannas. Similar to other tropical biomes, this biome has high species diversity. For example, the Everglades are home to some 11,000 species of seed-bearing plants, 25 species of orchids, 300 bird species, and 150 species of fish.
Tropical and Subtropical Moist Broadleaf Forests
The tropical and subtropical moist
broadleaf forests biome (also called moist tropical rain forest) occurs in a
zone about 10° of latitude either side of the equator. Annual rainfall
generally exceeds 250 centimeters (100 inches) and is evenly distributed
throughout the year. Temperature and humidity are relatively high through the
year. Flora is highly diverse: a typical hectare (2.5 acres) may contain as
many as 300 different tree species as compared to 20 to 30 in the temperate
zone. The various trees of the moist tropical rain forests are closely spaced
together and form a thick continuous canopy some 25 to 35 meters (80 to 115
feet) tall. Every so often this canopy is interrupted by the presence of very
tall emergent trees (up to 40 meters or 130 feet) that have wide buttressed
bases for support. Epiphytic orchids and bromeliads, as well as vines (lianas),
are very characteristic of the moist tropical rain forest biome. Some other
common plant species include ferns and palms. Most plants are evergreen with
large, dark green, leathery leaves. The ground surface of the moist tropical rain forest tends to be dark with only about 1% of the light intensity found above the forest canopy. These light poor conditions cause the understory to be sparsely vegetated. The few plants that grow at ground level do so by being able to tolerate low light levels. The moist tropical rain forest is also home to a great variety of animals. Some scientists believe that 30 to 50% of all of the Earth's animal species may be found in this biome. Most of these organisms are insects.
Decomposition is rapid in the tropical rain forest because of high temperatures and abundant moisture. Because of the frequent and intense rains, tropical soils are subject to extreme chemical weathering and leaching. These environmental conditions make tropical soils acidic and nutrient poor.
Tropical and Subtropical Dry Broadleaf Forests
Tropical and subtropical dry forests
(also called seasonal tropical forest or tropical dry forest) are found in
southern Mexico, southeastern Africa, central India, Indochina, Madagascar, New
Caledonia, eastern Bolivia, central Brazil, the Caribbean, and along the coasts
of Peru and Ecuador.This biome exists as a zone that borders the tropical and
subtropical moist broadleaf forests biome. Because of its geographical
location, the tropical and subtropical dry forest experiences a dry season that
lasts several months. This abiotic condition has a great effect on living
things in this biome. Many of these species that live here have specific
adaptations to help them survive the dry period. Consequently, deciduous trees
like teak, mahogany, and mountain ebony dominate these forests. During the
seasonal drought these trees loose their leaves to conserve water.The leafless
condition also causes more sunlight to reach ground surface and this condition
facilitates the growth of thick shrub layer. While less diverse than tropical
rain forests, seasonal tropical forests still have a vast assortment of
organisms.
Tropical and Subtropical Coniferous Forests
The tropical and subtropical
coniferous forests biome is characterized by diverse species of conifer
(needle-leaf) trees.This biome has a very limited distribution and is found
mainly in Mexico, Central America, and on the islands of Cuba, Dominican
Republic, and Haiti where low levels of precipitation and moderate temperature
variability occurs. The needle-leaf form of these trees is an adaptation to
drought. This biome shares some of the plant and animal species common to
tropical and subtropical savanna, dry broadleaf forest, and moist broadleaf
forest. Understory vegetation composed of shrubs and small trees is well
developed and diverse. Finally, many species of migratory birds and butterflies
spend their winter in this biome.
The Importance and
Conservation of Biomes
Because
we share the world with many other species of plants and animals, we must
consider the consequences of our actions. Over the past several decades,
increasing human activity has rapidly destroyed or polluted many ecological habitats
throughout the world. It is important to preserve all types of biomes as each
houses many unique forms of life. However, the continued heavy exploitation of
certain biomes, such as the forest and aquatic, may have more severe
implications.Conservation of Biomes
Forests are important as they are home to the most diverse biotic communties in the world. Hidden within these biomes are potential medicines and many thousands of unseen and undiscovered species. Also, forests have a global climate-buffering capacity, so their destruction may cause large-scale changes in global climate.
Logging has depleted many old-growth temperate forests. The increased demand for homes, paper, and other wood products have not allowed for much conservation. More recently, people have begun to realize that logging has cleared much of these forests. Wiser use of the forests and efforts to replant trees have helped to slow down the depletion of these communities.
Tropical forests have fallen victim to timber exploitation, slash and burn farming, and clearfelling for industrial use or cattle ranching, particularly in Latin America. Our increasing demand for meat products has spurred these events. For years, this destruction was occuring at a rapid rate. Over half of the world’s original tropical forests are already gone. Public attention to this exploitation have helped to alleviate the problem somewhat, though many challenges are still to be faced.
Aquatic biomes are probably the most important of all the biomes. Their medium, water, is a major natural resource. Water is the basis of life, it supports life, and countless species live in it for all or part of their lives. Freshwater biomes supply us with our drinking water and water for crop irrigation. The world’s oceans have an even greater effect on global climate than forests do. Water has a high capacity for heat, and because the Earth is mostly covered with water, the temperature of the atmosphere is kept fairly constant and able to support life. In addition to this climate-buffering capacity, the oceans contain several billion photosynthetic plankton which account for most of the photosynthesis occuring on Earth. Without these, there might not be enough oxygen to support such a large world population and complex animal life.
Freshwater biomes have suffered mainly from pollution. Runoff containing fertilizer and other wastes and industrial dumpings enter into rivers, ponds, and lakes and tend to promote abnormally rapid algae growth. When these algae die, dead organic matter accumulates in the water. This makes the water unusable and it kills many of the organisms living in the habitat. Stricter laws have helped to slow down this thoughtless pollution.
Overfishing and pollution have threatened to make oceans into ecological disaster areas. Industrial pollutants that are dumped upstream of estuaries have rendered many marine habitats unsuitable for life. Again, tighter regulations have been used to prevent further destruction of the ocean biomes.
By educating people about the consequences of our actions, we can all gain a better understanding of how to preserve the earth’s natural biomes. The areas that have been destroyed the most will never regain their original forms, but conservation will help to keep them from getting worse.
SUMMARY
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| Biomes
are often named for their predominant vegetation, and all have certain
characteristics such as microorganisms, fungi, and animals which have adapted
to their particular environment. As well, biomes also grade into one
another, and do not usually have sharp boundaries. In fact, if the area
of intergradation is large enough, it may itself be recognized as a separate
biome. In biomes of the same type, but located in different areas, the species found there may vary. For example, in the coniferous forests of North America, red spruce is common in the east, but is not found in other areas. There, black spruce and white spruce are abundant. Also, plants of different families are found in African and North American deserts, but the plants do resemble one another superficially. This similarity can arise due to convergent evolution, “the independant development of similarity between species as a result of their having similar ecological roles and selection pressures. Several communities may be represented in patches within a biome. For example, snowfall may break branches and small trees and cause openings in a coniferous forest, allowing deciduous species to grow. |










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